I'll take a crack at this, but Mike can do a better and more technical response.
The V1 USM (uncommanded servo movement) is a error that occurs when some bad data is accepted by the receiver. This can happen as the receiver is not fully checking the quality of the data in the entire packet. So a bad packet has a small chance to be accepted by the receiver. The packet data is basically arrives in a given order. For example, the first number would be channel one, the second number channel two, etc. There is nothing that states, channel one equals 1500, it just says 1500 in the correct place. This all works fine as long as the data is correct. If the receiver approves a bad packet of data, then you get random servo movements. With the Universal firmware there are additional checks in place to catch this bad data and reject, thus the issue that was already VERY rare is almost completely eliminated. But just in case a further measure was added that should a bad packet manage to pass both quality checks, the USM would only last 18ms as the firmware no longer accepts any data from the previous packet as being valid. (something V1 did to save on workload..?) So the next packet of data comes in, gets checked twice, etc.. With V1 I forget how long the USM could last, but it was a much longer time frame, and the risks of more bad packets being accepted was much higher.
I've probably done a horrible job and may not have the details fully correct here, but this is my understanding based on my memory. Sure mike will correct any major flaws.
The takeaway is that Universal has multiple improvements to fight USM making it the better choice. The other major addition is the AutoTune feature, for some receiver transmitter combinations will net noticeable range & packet performance. Basically AutoTune does a series of tests of the RF connection and adjusts the tune slightly and checks to see if the connection is better or worse, and it does this until it finds the sweet spot. Since all radio gear has some tolerances due to the components used, the frequencies can shift up or down slightly. The AutoTune adjusts for this to get the best signal from the transmitter. Just like tuning an old stereo system to remove the static from a music broadcast. (Younger people may not know what that was like, LOL.) To me this is possibly the most exciting improvement in this firmware.